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TO allow quantitative analysis of phylogeographic tive distribution in space) and the existence of a spatial data, Templeton et al.(1995) have developed a pattern sensu stricto (due to a spatial dependence of statistical framework called the nested cladistic analysis haplotype frequencies in a population to those of the (NCA) of geographical distance, which can be used to neighboring populations). infer past and ongoing processes using the genealogical We consider that both sources of genetic structure and geographical information available in population should be disentangled and tested successively. To test genetic surveys. It is the only available technique that the spatial distribution of haplotypes, it is important to incorporates the explicit testing of the geographic keep other aspects of the genetic structure constant, spread of a haplotype relative to its center or to the such as the level of fixation within populations …