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Human-driven declines of apex predators can trigger widespread impacts throughout ecological communities. Reduced apex predator occupancy or activity can release mesopredators from intraguild competition, with unknown repercussions on the ecological community. As exurban development continues to expand worldwide, it is important to document how mesopredators are impacted by the combined influences of apex predators and humans. We used motion-detecting camera traps to examine spatial and temporal patterns of meso- and apex predator occupancy and activity in a fragmented landscape in California. We hypothesized that both spatial and temporal partitioning among the carnivore guild would be affected by varied levels of human influence. We found that higher residential development reduced puma occupancy but was not related to the occupancy of mesopredators. Bobcats, grey foxes, and …