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CS Holling f, Bengt-Owe Jansson g, Simon Levin h Karl-GSran M~ iler i Charles Perrings J, David Pimentel k a Department of Economics, Stanford University, Stanford, CA 94305, USA b Department of Meteorology, University of Stockholm, 106 91 Stockholm, Sweden c Maryland International Institute for Ecological Economics, University of Maryland, Box 38, Solomons, MD 20688, USA a Faculty of Economics, Cambridge University, Cambridge CB3 9DD, UK e Beijer International Institute of Ecological Economics, Royal Swedish Academy of Sciences, Box 50005, S-104 05 Stockholm, Sweden f Department of Zoology, University of Florida, Gainesville, FL 32611, USA g Department of Systems Ecology, University of Stockholm, S-106 91 Stockholm, Sweden h Department of Ecology and Evolutionary Biology, Princeton University, Princeton, NJ 08544, USA i Beijer International Institute of Ecological Economics, Royal …