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Definitions and concepts of production ecology are presented as a basis for development of alternative production technologies characterized by their input-output combinations. With these concepts the relative importance of several growth factors and inputs is investigated to explain actual yield levels and resource-use efficiencies. Differences between potential and actual levels are analyzed to open ways for improved production technologies. The basis of the analysis is knowledge of basic physical, chemical, physiological and ecological processes at soil, field and crop level. New production technologies and their input-output combinations can be used in studies aimed at the exploration of options for sustainable agricultural production systems and land use. The concepts allow a systematic analysis and quantification of input-output combinations and clearly discriminate between bio-physical possibilities and …