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Freshwater midges, consisting of Chironomidae, Chaoboridae and Ceratopogonidae, were assessed as a biological proxy for palaeoclimate in eastern Beringia. The northwest North American training set consists of midge assemblages and data for 17 environmental variables collected from 145 lakes in Alaska, British Columbia, Yukon, Northwest Territories, and the Canadian Arctic Islands. Canonical correspondence analyses (CCA) revealed that mean July air temperature, lake depth, arctic tundra vegetation, alpine tundra vegetation, pH, dissolved organic carbon, lichen woodland vegetation and surface area contributed significantly to explaining midge distribution. Weighted averaging partial least squares (WA-PLS) was used to develop midge inference models for mean July air temperature (r _boot ²  = 0.818, RMSEP = 1.46°C), and transformed depth (ln (x+1 …