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Pollen and oceanographic data from deep ocean core MD95-2039 provide a centennial to millennial scale record of conditions offshore and of the vegetation of north-west Iberia for the period 10–65ka. The planktonic oxygen isotope record of this core, reflecting predominantly sea surface temperature (SST), shows a pattern of millennial-scale oscillations that is very similar to climatic changes recorded by the Greenland ice core records over the same interval. In turn, tree populations show a pattern of rapid expansions and contractions that follow the pronounced and abrupt isotopic shifts recorded offshore. Through Marine Isotope Stage (MIS) 3, this millennial-scale pattern of vegetation change, alternating between steppe and open woodland, is superimposed on a longer-term pattern of shrinking ericaceous heathland and decreasing size of successive interstadial tree populations. Trees persisted during the Last …