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This article first develops an integrated traffic-driving-networking simulator (ITDNS) intended for the design and evaluation of cyber transportation systems (CTS) and connected vehicle (CV) applications. The ITDNS allows a human driver to control a subject vehicle, in a virtual environment, that is capable of communicating with other vehicles and the infrastructure with CTS messages. The challenges associated with the integration of the three simulators, and how those challenges were overcome, are discussed. As an application example, an eco-signal system, which recommends the approach speed for vehicles approaching the intersection so as to minimize fuel consumption and emissions, was implemented in the ITDNS. Test drivers were then asked to virtually drive through a signalized corridor twice, one time with the eco-signal system in place and another without the system. Thanks to the human-in-the-loop …