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Dispersal is usually considered in terms of movement from one location and the consequences of arriving at other locations. Movement between these locations is often viewed as a linear process, on which the characteristics of the intervening area have little effect. The area between the beginning and ending of dispersal, however, is not a featureless matrix of unsuitable habitat, but is instead a richly textured mosaic of patches of different shapes, sizes, arrangements, and qualities: a landscape. Movement through such a mosaic is not likely to be linear, and how an individual traverses the landscape then becomes important. Different locations in a landscape are associated with different costs and benefits, and the movement of an individual determines, and is determined by, the combination of costs and benefits it encounters. The propositions that movement during dispersal may depend on the structure of landscapes, and that these landscape effects can influence population distribution and the probabilities of movement among locations, are supported by several studies that have used experimental model systems (EMSs) to document and model individual movements in fine-scale mosaics. These studies also illustrate the importance of the scales on which organisms perceive landscape structure. Assessing how landscapes may influence dispersal probabilities therefore requires a consideration of the interaction between the scale of landscape pattern and the scale on which organisms respond to that pattern. Landscapes change over time as a consequence of natural processes, such as succession, disturbance, or human land use. Habitat …