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Throughout 2014–2016, the California Current System (CCS) was characterized by large and persistent sea surface temperature anomalies (SSTa), which were accompanied by widespread ecological and socioeconomic consequences that have been documented extensively in the scientific literature and in the popular press. This marine heatwave and others have resulted in a heightened awareness of their potential impacts and prompted questions about if and when they may be predictable. Here, we use output from an ensemble of global climate forecast systems to document which aspects of the 2014–2016 CCS heatwave were predictable and how forecast skill, or lack thereof, relates to mechanisms driving the heatwave’s evolution. We focus on four prominent SSTa changes within the 2014–2016 period: (i) the initial onset of anomalous warming in early 2014, (ii) a second rapid SSTa increase in late 2014, (iii) a sharp reduction and subsequent return of warm SSTa in mid-2015, and (iv) another anomalous warming event in early 2016. Models exhibited clear forecast skill for the first and last of these fluctuations, but not the two in the middle. Taken together with the state of knowledge on the dominant forcing mechanisms of this heatwave, our results suggest that CCS SSTa forecast skill derives from predictable evolution of pre-existing SSTa to the west (as in early 2014) and the south (as in early 2016), while the inability of models to forecast wind-driven SSTa in late 2014 and mid-2015 is consistent with the lack of a moderate or strong El Niño or La Niña event preceding those periods. The multi-model mean forecast consistently …