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Reconstructing marine paleoenvironments from the skeletal hardparts of nektic organisms is often hampered by their unknown migrational pathways involving different water masses and their corresponding physico-chemical parameters. Despite significant progress over the last years, the reconstruction of migration patterns of extinct ammonoids is difficult because both, vertical and horizontal ones lead to an intricate combination of reconstructed seawater temperatures. Paleonvironmental data retrieved from exceptionally well-preserved lower Albian (Douvilleiceras mammillatum ammonite zone, uppermost CC8a nannofossil zone, 110.5–111.0 Ma) cephalopod shells from Madagascar (Mahajanga Basin, 40–42° southern latitude) are reconstructed based on a novel multi-proxy approach. Here we combine and contrast: (i) nacre tablet-paleobathymetry, (ii) Westermann Morphospace, and (iii) shell isotope …