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Twitter is one of the largest social networks using exclusively directed links among accounts. This makes the Twitter social graph much closer to the social graph supporting real life communications than, for instance, Facebook. Therefore, understanding the structure of the Twitter social graph is interesting not only for computer scientists, but also for researchers in other fields, such as sociologists. However, little is known about how the information propagation in Twitter is constrained by its inner structure. In this paper, we present an in-depth study of the macroscopic structure of the Twitter social graph unveiling the highways on which tweets propagate, the specific user activity associated with each component of this macroscopic structure, and the evolution of this macroscopic structure with time for the past 6 years. For this study, we crawled Twitter to retrieve all accounts and all social relationships (follow links …