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While the emergence of the Acheulean is well documented in East Africa at ~1.7 Ma, subsequent developments are less well understood and to some extent controversial. Here, we provide robust evidence regarding the time period between 1.6 Ma and 1.2 Ma, based on an interdisciplinary approach to the stratigraphic sequences exposed in the Gombore gully of Melka Kunture, in the upper Awash Valley of Ethiopia. Throughout the Pleistocene, the environment differed significantly from elsewhere in Africa because of the elevation at 2000 m asl, the cooler and rainy climate, the Afromontane vegetation, the development of endemic animal species, and the recurrent impact of volcanic activity. At Gombore IB, dated ~1.6 Ma, remains of Homo erectus/ergaster have been discovered, associated with a rich early Acheulean assemblage. The techno-typological analysis of the lithic record from Gombore Iγ (~1.4 Ma) and …